Abstract
The CCAAT boxes of the beta-like globin genes interact with three proteins: NF-Y, GATA-1 and NFE-6. We demonstrate that NFE-6 contains C/EBPgamma, and address its role in globin gene regulation by erythroid overexpression of C/EBPgamma, and a dominant-negative form C/EBPgammaDeltaB, in mice. Elevated levels of C/EBPgamma, but not C/EBPgammaDeltaB, increase expression of the (fetal) gamma-globin relative to the (adult) beta-globin gene. Interestingly, fetal liver erythropoiesis is ablated when the C/EBPgamma and C/EBPgammaDeltaB levels are further increased in homozygous transgenics. We suggest that targeted expression of dominant-negative leucine zipper proteins is a generally applicable approach to ablate specific tissues in mice.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Animals
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Base Sequence
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CCAAT-Binding Factor / genetics
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CCAAT-Enhancer-Binding Proteins / genetics*
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DNA Primers / genetics
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DNA-Binding Proteins / genetics
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Erythroid-Specific DNA-Binding Factors
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Erythropoiesis / genetics*
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Fetal Blood / cytology
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Fetal Blood / metabolism
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GATA1 Transcription Factor
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Gene Expression Regulation, Developmental
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Globins / genetics*
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Leucine Zippers / genetics
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Liver / embryology
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Mice
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Mice, Transgenic
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Multigene Family
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Transcription Factors / genetics
Substances
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CCAAT-Binding Factor
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CCAAT-Enhancer-Binding Proteins
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CCAAT-enhancer-binding protein-gamma
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DNA Primers
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DNA-Binding Proteins
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Erythroid-Specific DNA-Binding Factors
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GATA1 Transcription Factor
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Gata1 protein, mouse
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Transcription Factors
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Globins