Sites of positive allosteric modulation by neurosteroids on ionotropic gamma-aminobutyric acid receptor subunits

FEBS Lett. 2004 May 21;566(1-3):213-7. doi: 10.1016/j.febslet.2004.04.030.

Abstract

Neurosteroids are known as allosteric modulators of ionotropic gamma-aminobutyric acid (GABA) receptors. Here, we investigated sites of positive allosteric modulation by allotetrahydrodeoxycorticosterone (5alpha-THDOC) at GABA receptors using the technique of chimeragenesis and the Xenopus oocyte expression system. Our findings have demonstrated that the region from transmembrane segment (TM) 4 to the C-terminus of the GABA(A) receptor alpha1 subunit is crucial for the action of 5alpha-THDOC, but insufficient for the action of another neurosteroid allopregnanolone, suggesting that a specific region critical for neurosteroid action at GABA receptors exists in the domain between TM4 and the C-terminus of GABA receptor subunits.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Allosteric Regulation
  • Allosteric Site
  • Amino Acid Sequence
  • Animals
  • Conserved Sequence
  • Desoxycorticosterone / analogs & derivatives*
  • Desoxycorticosterone / pharmacology*
  • Dose-Response Relationship, Drug
  • Electrophysiology
  • Female
  • GABA-A Receptor Agonists*
  • Humans
  • Molecular Sequence Data
  • Oocytes / drug effects
  • Oocytes / metabolism
  • Pregnanolone / pharmacology*
  • Protein Subunits
  • Receptors, GABA-A / chemistry
  • Receptors, GABA-A / genetics
  • Recombinant Proteins / agonists
  • Recombinant Proteins / chemistry
  • Recombinant Proteins / genetics
  • Xenopus Proteins / agonists
  • Xenopus Proteins / chemistry
  • Xenopus Proteins / genetics
  • Xenopus laevis

Substances

  • GABA-A Receptor Agonists
  • Protein Subunits
  • Receptors, GABA-A
  • Recombinant Proteins
  • Xenopus Proteins
  • Desoxycorticosterone
  • tetrahydrodeoxycorticosterone
  • Pregnanolone