Prolactin modulates IL-8 production induced by porins or LPS through different signaling mechanisms

Immunobiology. 2004;209(7):523-33. doi: 10.1016/j.imbio.2004.06.001.

Abstract

Prolactin (PRL) induces cell proliferation and cell differentiation through the well-known mitogen-activated protein kinases (MAPKs) and Janus kinase (JAK)/signal transducer and activator of transcription (STAT) pathways, depending on the cell line. MAPKs play a central role in signaling transduction mechanisms that transmit mitogenic or differentiation signals from an activated receptor to the intracellular machinery. All of the cytokine receptors that activate the JAK/STAT pathway also activate the MAPK pathway. The aim of the present study was to delineate the signal pathways implicated in IL-8 release by THP-1 cells, pretreated with PRL, after stimulation with either lipopolysaccharide (LPS) or porins from Salmonella enterica serovar Typhimurium. PRL activates the JAK2/STAT1-3 signaling pathway, while LPS or porins from S. enterica serovar Typhimurium does not induce any phosphorylation of this pathway. However, in THP-1 cells, the combination of PRL followed by either S. enterica serovar Typhimurium LPS or porins produced a greater MEK1-MEK2/MAPKs activation response than treatment with PRL alone. Similarly, PRL pretreatment of THP-1 cells resulted in an increase in IL-8 release in response to stimulation with either LPS or porins. This additive effect on IL-8 release was reduced when the cells were also treated with PD-098059, a selective inhibitor of the MEK1 activator and the MAPK cascade, or SB203580, a specific inhibitor of the p38 pathway, or AG490, a specific JAK/STAT pathway inhibitor, providing evidence that there are different signal pathways activated which have a cumulative effect.

MeSH terms

  • Blotting, Western
  • Cell Line
  • Cytokines / metabolism
  • Electrophoresis, Polyacrylamide Gel
  • Enzyme-Linked Immunosorbent Assay
  • Humans
  • Interleukin-8 / metabolism*
  • L-Lactate Dehydrogenase / metabolism
  • Lipopolysaccharides / metabolism*
  • Mitogen-Activated Protein Kinases / metabolism
  • Porins / metabolism*
  • Prolactin / metabolism*
  • Protein-Tyrosine Kinases / metabolism
  • Salmonella typhimurium / metabolism
  • Signal Transduction / physiology*
  • Trans-Activators / metabolism

Substances

  • Cytokines
  • Interleukin-8
  • Lipopolysaccharides
  • Porins
  • Trans-Activators
  • Prolactin
  • L-Lactate Dehydrogenase
  • Protein-Tyrosine Kinases
  • Mitogen-Activated Protein Kinases