Morphological analysis of flowers was carried out in Paeonia cultivars. Some unusual alternations of floral organs were described: sepal-(petal-stamen) x N-carpel, where 2 < or = n < or = 4 (appearance of an additional zone of petal and stamen formation in the medial flower part). The identity of floral organs was not affected in the flowers with this unusual alternation. It was shown on the basis of mathematical simulation of the genes responsible for flower development that these alternations may be determined by increased pool of stem cells, which may lead to delayed termination of flower development.