Astrocytes express N-methyl-D-aspartate receptor subunits in development, ischemia and post-ischemia

Neurochem Res. 2010 Dec;35(12):2124-34. doi: 10.1007/s11064-010-0325-x. Epub 2010 Nov 30.

Abstract

The expression of the N-methyl-D-aspartate receptor (NMDA-R) in astrocytes is controversial. The receptor is commonly considered neuron-specific. We showed that astrocytes in primary cultures differentially expressed mRNA of NMDA-R subunits, NR1, NR2A and NR2B, in development, ischemia and post-ischemia. One-week-old cultures expressed detectable NR1 mRNA, which fell significantly at 2 weeks and became barely detectable at 4 weeks. NR2A and NR2B mRNA were both significantly up-regulated from 1 to 2 weeks. In 4 weeks, 2 h of ischemia caused a significant up-regulation of NR1 and NR2B mRNA; while 6 h caused down-regulation of NR2A mRNA. Under 3 h of post-ischemia, only NR1 mRNA was increased. Ischemia induced the expression of major NMDA-R effecter, nitric oxide synthase 1, which was unaffected by AMPA-R antagonist CNQX, but dose-dependently inhibited by NMDA-R specific antagonist MK-801. These findings reflected that astrocyte could express inducible functional NMDA receptors without the presence of neurons.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Astrocytes / metabolism*
  • Brain Ischemia / metabolism*
  • Brain Ischemia / pathology
  • Down-Regulation
  • Fluorescent Antibody Technique
  • Glutamic Acid / pharmacology
  • Hydrogen Peroxide / pharmacology
  • Mice
  • Mice, Inbred ICR
  • Nitric Oxide Synthase Type I / metabolism
  • RNA, Messenger / genetics
  • Receptors, N-Methyl-D-Aspartate / genetics
  • Receptors, N-Methyl-D-Aspartate / metabolism*
  • Up-Regulation

Substances

  • RNA, Messenger
  • Receptors, N-Methyl-D-Aspartate
  • Glutamic Acid
  • Hydrogen Peroxide
  • Nitric Oxide Synthase Type I
  • Nos1 protein, mouse