Evaluation of the effects of erythritol on gene expression in Brucella abortus

PLoS One. 2012;7(12):e50876. doi: 10.1371/journal.pone.0050876. Epub 2012 Dec 14.

Abstract

Bacteria of the genus Brucella have the unusual capability to catabolize erythritol and this property has been associated with their virulence mainly because of the presence of erythritol in bovine foetal tissues and because the attenuated S19 vaccine strain is the only Brucella strain unable to oxydize erythritol. In this work we have analyzed the transcriptional changes produced in Brucella by erythritol by means of two high throughput approaches: RNA hybridization against a microarray containing most of Brucella ORF's constructed from the Brucella ORFeome and next generation sequencing of Brucella mRNA in an Illumina GAIIx platform. The results obtained showed the overexpression of a group of genes, many of them in a single cluster around the ery operon, able to co-ordinately mediate the transport and degradation of erythritol into three carbon atoms intermediates that will be then converted into fructose-6P (F6P) by gluconeogenesis. Other induced genes participating in the nonoxidative branch of the pentose phosphate shunt and the TCA may collaborate with the ery genes to conform an efficient degradation of sugars by this route. On the other hand, several routes of amino acid and nucleotide biosynthesis are up-regulated whilst amino acid transport and catabolism genes are down-regulated. These results corroborate previous descriptions indicating that in the presence of erythritol, this sugar was used preferentially over other compounds and provides a neat explanation of the the reported stimulation of growth induced by erythritol.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Amino Acids / metabolism
  • Animals
  • Brucella abortus / drug effects*
  • Brucella abortus / metabolism
  • Carbohydrates / chemistry
  • Cattle
  • Cluster Analysis
  • Erythritol / pharmacology*
  • Fructosephosphates / metabolism
  • Gene Expression Regulation, Bacterial / drug effects*
  • Genome, Bacterial
  • Models, Biological
  • Nucleotides / chemistry
  • Oligonucleotide Array Sequence Analysis
  • Open Reading Frames
  • RNA, Messenger / metabolism
  • Real-Time Polymerase Chain Reaction / methods
  • Sequence Analysis, DNA
  • Sequence Analysis, RNA
  • Virulence

Substances

  • Amino Acids
  • Carbohydrates
  • Fructosephosphates
  • Nucleotides
  • RNA, Messenger
  • fructose-6-phosphate
  • Erythritol

Grants and funding

This work was supported by the Ministerio de Ciencia y Tecnología of Spain (BIO2005-04985 and AGL2008-04514 to ILG, and BIO2007-63656 and BFU2011-25658 to FJS) and Instituto de Salud Carlos III (PI050894 to JMG-L). Fellowships support to CV from the Gobierno Vasco and to MCR from Fundación Marqués de Valdecilla-IFIMAV are gratefully acknowledged. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.