A multilocus species delimitation reveals a striking number of species of coralline algae forming Maerl in the OSPAR maritime area

PLoS One. 2014 Aug 11;9(8):e104073. doi: 10.1371/journal.pone.0104073. eCollection 2014.

Abstract

Maerl beds are sensitive biogenic habitats built by an accumulation of loose-lying, non-geniculate coralline algae. While these habitats are considered hot-spots of marine biodiversity, the number and distribution of maerl-forming species is uncertain because homoplasy and plasticity of morphological characters are common. As a result, species discrimination based on morphological features is notoriously challenging, making these coralline algae the ideal candidates for a DNA barcoding study. Here, mitochondrial (COI-5P DNA barcode fragment) and plastidial (psbA gene) sequence data were used in a two-step approach to delimit species in 224 collections of maerl sampled from Svalbard (78°96'N) to the Canary Islands (28°64'N) that represented 10 morphospecies from four genera and two families. First, the COI-5P dataset was analyzed with two methods based on distinct criteria (ABGD and GMYC) to delineate 16 primary species hypotheses (PSHs) arranged into four major lineages. Second, chloroplast (psbA) sequence data served to consolidate these PSHs into 13 secondary species hypotheses (SSHs) that showed biologically plausible ranges. Using several lines of evidence (e.g. morphological characters, known species distributions, sequences from type and topotype material), six SSHs were assigned to available species names that included the geographically widespread Phymatolithon calcareum, Lithothamnion corallioides, and L. glaciale; possible identities of other SSHs are discussed. Concordance between SSHs and morphospecies was minimal, highlighting the convenience of DNA barcoding for an accurate identification of maerl specimens. Our survey indicated that a majority of maerl forming species have small distribution ranges and revealed a gradual replacement of species with latitude.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Atlantic Ocean
  • Conservation of Natural Resources*
  • DNA Barcoding, Taxonomic
  • Genetic Loci / genetics*
  • Phylogeny
  • Rhodophyta / classification*
  • Rhodophyta / genetics*

Grants and funding

This work was funded by Ministerio de Ciencia e Innovación of Spain (grant CTM2010-18787, partially co-founded by FEDER, Fondo Europeo de Desarrollo Regional), and Xunta de Galicia (grant 10MMA103003PR). Samples from French Brittany were collected with the support of a European Community-Research Infrastructure Action under the FP7 “Capacities” Specific Programme (ASSEMBLE grant agreement no. 227799). Several authors were supported by fellowships: CP (Xunta de Galicia Axudas á etapa predoutoral do Plan I2C-2011, and Spain’s Ministerio de Educación Programa FPU-2010), LL (Universidade da Coruña Contratos Predoutorais-2012), and VP (Xunta de Galicia Axudas posdoutorais do Plan I2C-2012). VP acknowledges support for examining type material (Action Transversale du Muséum National d’Histoire Naturelle, project "Taxonomie moléculaire: DNA Barcode et gestion durable des collections"). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.