During Drosophila disc regeneration, JAK/STAT coordinates cell proliferation with Dilp8-mediated developmental delay

Proc Natl Acad Sci U S A. 2015 May 5;112(18):E2327-36. doi: 10.1073/pnas.1423074112. Epub 2015 Apr 20.

Abstract

Regeneration of fragmented Drosophila imaginal discs occurs in an epimorphic manner involving local cell proliferation at the wound site. After disc fragmentation, cells at the wound site activate a restoration program through wound healing, regenerative cell proliferation, and repatterning of the tissue. However, the interplay of signaling cascades driving these early reprogramming steps is not well-understood. Here, we profiled the transcriptome of regenerating cells in the early phase within 24 h after wounding. We found that JAK/STAT signaling becomes activated at the wound site and promotes regenerative cell proliferation in cooperation with Wingless (Wg) signaling. In addition, we showed that the expression of Drosophila insulin-like peptide 8 (dilp8), which encodes a paracrine peptide to delay the onset of pupariation, is controlled by JAK/STAT signaling in early regenerating discs. Our findings suggest that JAK/STAT signaling plays a pivotal role in coordinating regenerative disc growth with organismal developmental timing.

Keywords: Drosophila imaginal discs; JAK/STAT signaling; epimorphic regeneration.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Animals
  • Body Patterning
  • Cell Lineage
  • Cell Proliferation
  • Cluster Analysis
  • Drosophila / physiology*
  • Drosophila Proteins / metabolism*
  • Gene Expression Regulation
  • Imaginal Discs / physiology*
  • Intercellular Signaling Peptides and Proteins / metabolism*
  • Janus Kinases / metabolism
  • Oligonucleotide Array Sequence Analysis
  • Principal Component Analysis
  • Regeneration
  • STAT Transcription Factors / metabolism*
  • Signal Transduction
  • Transcription Factors / metabolism
  • Transcriptome
  • Wound Healing*

Substances

  • Drosophila Proteins
  • Intercellular Signaling Peptides and Proteins
  • STAT Transcription Factors
  • Transcription Factors
  • insulin-like peptide 8, Drosophila
  • Janus Kinases
  • hop protein, Drosophila

Associated data

  • GEO/GSE54868