1 Department of Genetics, Yale School of Medicine, New Haven, Connecticut, United States of America; Department of Molecular Biophysics and Biochemistry, Yale University, New Haven, Connecticut, United States of America; Department of Therapeutic Radiology, Yale School of Medicine, New Haven, Connecticut, United States of America; Yale Cancer Center, New Haven, Connecticut, United States of America.
2 Department of Cell Biology, Yale School of Medicine, New Haven, Connecticut, United States of America.
3 Department of Genetics, Yale School of Medicine, New Haven, Connecticut, United States of America.