The Presence of Clitoromegaly in the Nonclassical Form of 21-Hydroxylase Deficiency Could Be Partially Modulated by the CAG Polymorphic Tract of the Androgen Receptor Gene

PLoS One. 2016 Feb 5;11(2):e0148548. doi: 10.1371/journal.pone.0148548. eCollection 2016.

Abstract

Background: In the nonclassical form (NC), good correlation has been observed between genotypes and 17OH-progesterone (17-OHP) levels. However, this correlation was not identified with regard to the severity of hyperandrogenic manifestations, which could depend on interindividual variability in peripheral androgen sensitivity. Androgen action is modulated by the polymorphic CAG tract (nCAG) of the androgen receptor (AR) gene and by polymorphisms in 5α-reductase type 2 (SRD5A2) enzyme, both of which are involved in the severity of hyperandrogenic disorders.

Objectives: To analyze whether nCAG-AR and SRD5A2 polymorphisms influence the severity of the nonclassical phenotype.

Patients: NC patients (n = 114) diagnosed by stimulated-17OHP ≥10 ng/mL were divided into groups according to the beginning of hyperandrogenic manifestations (pediatric and adolescent/adult) and CYP21A2 genotypes (C/C: homozygosis for mild mutations; A/C: compound heterozygosis for severe/mild mutations).

Methods: CYP21A2 mutations were screened by allelic-specific PCR, MLPA and/or sequencing. HpaII-digested and HpaII-undigested DNA samples underwent GeneScan analysis to study nCAG, and the SRD5A2 polymorphisms were screened by RLFP.

Results: Mean nCAG did not differ among pediatric, adolescent/adult and asymptomatic subjects. In the C/C genotype, we observed a significantly lower frequency of longer CAG alleles in pediatric patients than in adolescent/adults (p = 0.01). In patients carrying the A/C genotype, the frequencies of shorter and longer CAG alleles did not differ between pediatric patients and adolescent/adults (p>0.05). Patients with clitoromegaly had significantly lower weighted CAG biallelic mean than those without it: 19.1±2.7 and 21.6±2.5, respectively (p = 0.007), independent of the CYP21A2 genotype's severity. The SRD5A2 polymorphisms were not associated with the variability of hyperandrogenic NC phenotypes.

Conclusions: In this series, we observed a modulatory effect of the CAG-AR tract on clinical manifestations of the NC form. Although the NC form is a monogenic disorder, our preliminary data suggested that the interindividual variability of the hyperandrogenic phenotype could arise from polygenic interactions.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • 3-Oxo-5-alpha-Steroid 4-Dehydrogenase / genetics
  • Adolescent
  • Adrenal Hyperplasia, Congenital / diagnosis
  • Adrenal Hyperplasia, Congenital / genetics*
  • Adrenal Hyperplasia, Congenital / pathology*
  • Adult
  • Alleles
  • Child
  • Child, Preschool
  • Clitoris / pathology*
  • Female
  • Gene Frequency
  • Genetic Association Studies*
  • Genotype
  • Humans
  • Male
  • Membrane Proteins / genetics
  • Mutation
  • Phenotype
  • Polymorphism, Genetic*
  • Receptors, Androgen / genetics*
  • Severity of Illness Index
  • Steroid 21-Hydroxylase / blood
  • Steroid 21-Hydroxylase / genetics
  • Steroid 21-Hydroxylase / metabolism
  • Trinucleotide Repeat Expansion
  • Trinucleotide Repeats*
  • X Chromosome Inactivation
  • Young Adult

Substances

  • Membrane Proteins
  • Receptors, Androgen
  • CYP21A2 protein, human
  • Steroid 21-Hydroxylase
  • 3-Oxo-5-alpha-Steroid 4-Dehydrogenase
  • SRD5A2 protein, human

Supplementary concepts

  • Congenital adrenal hyperplasia due to 21 hydroxylase deficiency

Grants and funding

This research was supported by grants from the Sao Paulo Research Foundation (FAPESP) #2014/07878-4, to Moura-Massari V by FAPESP #08/51624-6 and FAPESP #05/04726-0, and to Bachega TASS and Mendonca BB by Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) #308318/2012-9 and 305743/2011-2, respectively. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.