1 Nottingham Digestive Diseases Centre, University of Nottingham, Nottingham, UK.
2 Derby Teaching Hospitals NHS Foundation Trust, Nottingham, UK.
3 Northern General Hospital, Sheffield, UK.
4 The NIHR Nottingham Biomedical Research Centre at Nottingham University Hospitals NHS Trust and University of Nottingham, Nottingham Digestive Diseases Centre, School of Medicine, University of Nottingham, Nottingham, UK.
5 NIHR Nottingham Biomedical Research Centre, Nottingham, UK.
6 Human Molecular Genetics, University of Heidelberg, Heidelberg, Germany.
7 Department of Therapeutics and Molecular Medicine, University of Nottingham, Nottingham, UK.