1 School of Biological Sciences, The University of Hong Kong, Pokfulam, Hong Kong.
2 School of Biological Sciences, The University of Hong Kong, Pokfulam, Hong Kong; Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544, USA.
3 Environmental Futures Research Institute and School of Environment and Science, Griffith University, Nathan, QLD, Australia.
4 School of Biological Sciences, The University of Hong Kong, Pokfulam, Hong Kong; Environmental Futures Research Institute and School of Environment and Science, Griffith University, Nathan, QLD, Australia. Electronic address: lashton@hku.hk.