1 Department of Biological Sciences, Virginia Tech, Blacksburg, VA, 24061, USA.
2 Newe Ya'ar Research Center, 30095, Ramat, Yishay, Israel.
3 United States Department of Agriculture, Agricultural Research Service, and Department of Horticulture, University of Wisconsin, Madison, WI, 53706, USA.
4 Department of Biological Sciences, Virginia Tech, Blacksburg, VA, 24061, USA. tholl@vt.edu.