1 Tobacco Research Institute, Chinese Academy of Agricultural Sciences, Qingdao, China.
2 Key Laboratory of Horticultural Plant Biology of Ministry of Education, College of Horticulture and Forestry Science, Huazhong Agricultural University, Wuhan, China.
3 Biotechnology Research Institute, Chinese Academy of Agricultural Sciences, Beijing, China.
4 Department of Plant Biotechnology and Bioinformatics, Ghent University, Ghent, Belgium.
5 VIB Center for Plant Systems Biology, Ghent, Belgium.