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The doubly phosphorylated form of HPr, HPr(Ser~P)(His-P), is abundant in exponentially growing cells of Streptococcus thermophilus and phosphorylates the lactose transporter LacS as efficiently as HPr(His~P).
Cochu A, Roy D, Vaillancourt K, Lemay JD, Casabon I, Frenette M, Moineau S, Vadeboncoeur C. Cochu A, et al. Appl Environ Microbiol. 2005 Mar;71(3):1364-72. doi: 10.1128/AEM.71.3.1364-1372.2005. Appl Environ Microbiol. 2005. PMID: 15746339 Free PMC article.